Plants of the Gila Wilderness
Presented in Association with the
Western New Mexico University Department
of Natural Sciences
Frullania cleistostoma/saxicola complex
Family: Frullaniaceae
Status: Native
Synonyms:
Awaits phylogenetic treatment
The genus Frullania has been re-evaluated for the state of New Mexico by John Atwood of the Missouri Botanical Garden.
Frullania inflata, formerly the most diagnosed species of the genus in New Mexico, is no longer thought to exist in this state. The specimens that were formerly assigned to Frullania inflata are now thought to mostly be part of an as yet poorly defined complex of more than one species-- currently referred to as the "cleistostoma/saxicola complex." A general description of the cleistostoma/saxicola complex as represented by this specimen annotated to be such by John Atwood is-- "Plants complicate-bilobed, medium sized (0.8-1.3mm), dark green to brownish-green; lobe is orbicular; lobule is explanate and less than half the size of the lobe; underleaves biobed, bifid to one-half; slightly larger than the stem, margins entire; leaf cells with smooth cell walls; oil bodies 3 to 5 per cell; monoicious; perianths occur on short or long branches, habitat on bark or rocks." (thanks John Atwood & Karen Blisard!)
Many thanks to the Missouri Botanical Garden and John Atwood for all their help and loaning this specimen for photography.
Please click on an image for a larger file.
Frullania cleistostoma, collection L.R. Stark & R. Castetter (as Frullania inflata, with annotation by John Atwood to F. cleistostoma, 2020), New Mexico, Doña Ana County, Organ Mountains, E side slopes and
ravines, between Sugarloaf Peak and Aguirre Springs Recreation Site, near Indian Hollow, on shaded
rock, July 12, 1980; photos by Russ Kleinman, Karen Blisard & Kelly Allred, December 1, 2023
Frullania cleistostoma, photomicrograph of stems with 2 perianths, collection L.R. Stark & R. Castetter (as Frullania inflata, with annotation by John Atwood to F. cleistostoma, 2020), New Mexico, Doña Ana County, Organ Mountains, E side slopes and
ravines, between Sugarloaf Peak and Aguirre Springs Recreation Site, near Indian Hollow, on shaded
rock, July 12, 1980; photos by Russ Kleinman, Karen Blisard & Kelly Allred, December 1, 2023
Frullania cleistostoma, photomicrograph of orbicular leaves, collection L.R. Stark & R. Castetter (as Frullania inflata, with annotation by John Atwood to F. cleistostoma, 2020), New Mexico, Doña Ana County, Organ Mountains, E side slopes and
ravines, between Sugarloaf Peak and Aguirre Springs Recreation Site, near Indian Hollow, on shaded
rock, July 12, 1980; photos by Russ Kleinman, Karen Blisard & Kelly Allred, December 1, 2023
Frullania cleistostoma, photomicrograph of lobules and underleaves, collection L.R. Stark & R. Castetter (as Frullania inflata, with annotation by John Atwood to F. cleistostoma, 2020), New Mexico, Doña Ana County, Organ Mountains, E side slopes and
ravines, between Sugarloaf Peak and Aguirre Springs Recreation Site, near Indian Hollow, on shaded
rock, July 12, 1980; photos by Russ Kleinman, Karen Blisard & Kelly Allred, December 1, 2023
Frullania cleistostoma, photomicrograph of underleaf & lobule, collection L.R. Stark & R. Castetter (as Frullania inflata, with annotation by John Atwood to F. cleistostoma, 2020), New Mexico, Doña Ana County, Organ Mountains, E side slopes and
ravines, between Sugarloaf Peak and Aguirre Springs Recreation Site, near Indian Hollow, on shaded
rock, July 12, 1980; photos by Russ Kleinman, Karen Blisard & Kelly Allred, December 1, 2023
Frullania cleistostoma, photomicrograph of underleaf and lobule, collection L.R. Stark & R. Castetter (as Frullania inflata, with annotation by John Atwood to F. cleistostoma, 2020), New Mexico, Doña Ana County, Organ Mountains, E side slopes and
ravines, between Sugarloaf Peak and Aguirre Springs Recreation Site, near Indian Hollow, on shaded
rock, July 12, 1980; photos by Russ Kleinman, Karen Blisard & Kelly Allred, December 1, 2023
Frullania cleistostoma, photomicrograph of relationship of lobule to lobe, collection L.R. Stark & R. Castetter (as Frullania inflata, with annotation by John Atwood to F. cleistostoma, 2020), New Mexico, Doña Ana County, Organ Mountains, E side slopes and
ravines, between Sugarloaf Peak and Aguirre Springs Recreation Site, near Indian Hollow, on shaded
rock, July 12, 1980; photos by Russ Kleinman, Karen Blisard & Kelly Allred, December 1, 2023
Frullania cleistostoma, photomicrograph of cells of lobe, collection L.R. Stark & R. Castetter (as Frullania inflata, with annotation by John Atwood to F. cleistostoma, 2020), New Mexico, Doña Ana County, Organ Mountains, E side slopes and
ravines, between Sugarloaf Peak and Aguirre Springs Recreation Site, near Indian Hollow, on shaded
rock, July 12, 1980; photos by Russ Kleinman, Karen Blisard & Kelly Allred, December 1, 2023
Back to the Index