Plants of the Gila Wilderness
Presented in Association with the
Western New Mexico University Department
of Natural Sciences
Grimmia torquata Drummond
Family: Grimmiaceae
Status: Native
Synonyms:
Grimmia pellucida Kindberg
Grimmia prolifera Müller Hal. & Kindberg
Grimmia pseudotorquata Kindberg
Grimmia tortifolia Kindberg
Grimmia torquata should be fairly easy to recognize by virtue of its
curly-Q leaves when dry, but especially because of the abundant multicellular red-brown
globular, stalked gemmae present at the base of the abaxial (back) surfaces of some of the apical leaves
on stems. Many of the leaves have the stalks remaining after the gemmae have dispersed,
deceiving one into initially thinking that the stalks themselves represent the gemmae.
Many thanks to the University of Colorado Herbarium (COLO), Erin Tripp & Dina Clark for permitting us
to photograph this specimen.
Please click on an image for a larger file.
Grimmia torquata, collection and ID by R.C. Wittmann, Colorado, Boulder County,
Rocky Mountains National Park, Allens Park Quadrangle, Wild Basin along Thunder Lake trail, July 26,
2003; photos by Russ Kleinman, Karen Blisard, Kelly Allred, Stacey Anderson & Ron Wittmann,
January 19, 2023
Grimmia torquata, photomicrograph of twisted and contorted stems, collection and ID by R.C. Wittmann, Colorado, Boulder County,
Rocky Mountains National Park, Allens Park Quadrangle, Wild Basin along Thunder Lake trail, July 26,
2003; photos by Russ Kleinman, Karen Blisard, Kelly Allred, Stacey Anderson & Ron Wittmann,
January 19, 2023
Grimmia torquata, photomicrograph of leaf (gemma adhering to leaf, but didn't originate
in current position since not on costa), collection and ID by R.C. Wittmann, Colorado, Boulder County,
Rocky Mountains National Park, Allens Park Quadrangle, Wild Basin along Thunder Lake trail, July 26,
2003; photos by Russ Kleinman, Karen Blisard, Kelly Allred, Stacey Anderson & Ron Wittmann,
January 19, 2023
Grimmia torquata, photomicrograph of leaf (gemma have dispersed, only stalks remain), collection and ID by R.C. Wittmann, Colorado, Boulder County,
Rocky Mountains National Park, Allens Park Quadrangle, Wild Basin along Thunder Lake trail, July 26,
2003; photos by Russ Kleinman, Karen Blisard, Kelly Allred, Stacey Anderson & Ron Wittmann,
January 19, 2023
Grimmia torquata, photomicrograph of leaf (gemmae on stalks in proper position at base
of leaf on abaxial surface), collection and ID by R.C. Wittmann, Colorado, Boulder County,
Rocky Mountains National Park, Allens Park Quadrangle, Wild Basin along Thunder Lake trail, July 26,
2003; photos by Russ Kleinman, Karen Blisard, Kelly Allred, Stacey Anderson & Ron Wittmann,
January 19, 2023
Grimmia torquata, photomicrograph of gemmae on abaxial costal surface at leaf base, collection and ID by R.C. Wittmann, Colorado, Boulder County,
Rocky Mountains National Park, Allens Park Quadrangle, Wild Basin along Thunder Lake trail, July 26,
2003; photos by Russ Kleinman, Karen Blisard, Kelly Allred, Stacey Anderson & Ron Wittmann,
January 19, 2023
Grimmia torquata, photomicrograph of leaf apex, collection and ID by R.C. Wittmann, Colorado, Boulder County,
Rocky Mountains National Park, Allens Park Quadrangle, Wild Basin along Thunder Lake trail, July 26,
2003; photos by Russ Kleinman, Karen Blisard, Kelly Allred, Stacey Anderson & Ron Wittmann,
January 19, 2023
Grimmia torquata, photomicrograph of dispersing gemmae, collection and ID by R.C. Wittmann, Colorado, Boulder County,
Rocky Mountains National Park, Allens Park Quadrangle, Wild Basin along Thunder Lake trail, July 26,
2003; photos by Russ Kleinman, Karen Blisard, Kelly Allred, Stacey Anderson & Ron Wittmann,
January 19, 2023
Grimmia torquata, photomicrograph of leaf cross section, collection and ID by R.C. Wittmann, Colorado, Boulder County,
Rocky Mountains National Park, Allens Park Quadrangle, Wild Basin along Thunder Lake trail, July 26,
2003; photos by Russ Kleinman, Karen Blisard, Kelly Allred, Stacey Anderson & Ron Wittmann,
January 19, 2023
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